Druon ignotum (Bassett, 1881), comb. nov.

Cynips ignota Bassett, 1881: 106, female, gall.
Andricus ignota (Bassett): Ashmead, 1885: 295.
Dryophanta ignota (Bassett): Ashmead, 1887: 127.
Diplolepis ignota (Bassett): Dalla Torre & Kieffer, 1910: 360. [NOT Diplolepis ignota Osten Sacken, 1863]
Rhodites ignota (Bassett): Dalla Torre, 1893: 127. [NOT Andricus ignota Bassett, 1900 (junior homonym).]
Diplolepis ignota (Bassett): Weld, 1926: 26.
Andricus ignotus (Bassett): Weld, 1951: 634.

Gall. Sexual galls (Fig. 100) occur singly or in groups in buds of Q. macrocarpa. They are apparently undifferentiated ovoid cells approximately 2 mm long, integral within buds and young growing shoots, and cause little or no visible external deformity or damage. Galls are generally only detectable by the emergence holes left by the adult insects.

Biology. Alternate asexual and sexual generations are known (see comments). Asexual galls have been recorded on leaves of Q. bicolor and Q. macrocarpa (Section Quercus, Series Prinoideae), mature in October–November, adult females emerge very early the following spring (late March to late April in Edmonton, Alberta, Canada), often on the first warm days after accumulated snow has melted and fallen leaves bearing galls are exposed. Emerged females fly up to buds and oviposit. The sexual generation adults emerged in Edmonton in early June. Females were observed ovipositing multiply along the midrib on the upper leaf surface in a zipper-like pattern, suggesting that asexual females in a given gall cluster are sisters. Galls of the asexual generation first became apparent on leaves in Edmonton in late July.

Distribution. USA: Massachusetts, Connecticut, Illinois, Minnesota (Burks 1979), Nebraska (Weld 1926); Canada: Alberta, Manitoba.