Phylloxera intermedia Pergande

Galls on leaves of Carya alba (= ovata) are circular, reddish or yellowish, 3-6 mm in diameter, convex to conical on upper side of leaf without any trace of a central nipple, paler and almost flat with a small closed nipple on underside. Alatae emerge in late may-June, mainly from a jagged opening formed by splitting of the upper surface of the gall (Pergande 1904). Locality not stated; probably in vicinity of Washington, D.C. The life cycle is unknown.

Phylloxera intermedia

Carya sp.
Conical or globose leaf galls with distinct orifice and containing aphids
Galls smaller though similar to P picta, opening on upper surface, more convex and often with a slight depression above, no opening above before maturity, a minute, closed nipple beneath, diameter 3 to 6 mm, height 2 to 3 mm.

Phylloxera intermedia Pergande, n. sp.

The gall of this species appears to be intermediate between Ph. c. -septum and Ph. picta. Its inner construction resembles that of c. -septum on account of the presence of a rudiment of the central membrane, though its general form, especially in the larger specimens, is more like that of picta, from which, however, it differs in the much thicker walls and the absence of a nipple on the upper side.

The transverse diameter of this gall varies between 3 and 6 mm and its vertical diameter between 2 and 3 mm. The shape above is regularly convex in the larger galls and slightly conical in the smaller ones, and without a trace of a nipple or opening before maturity. On the under side it projects but slightly and is provided with a minute, but closed nipple, the insect issuing from the upper surface which splits in a more or less jagged, round or oval opening, the bracts of which curve more or less outward. In some few cases the nipple on the under side also opens sufficiently to allow the insects to emerge from both sides. The color of the galls above is reddish or yellowish, generally surrounded by a paler ring, and beneath paler than the leaf. It occurs on Hicoria alba [Carya tomentosa] in company with the other two, and matures from the end of May to the middle of June.

Phylloxera intermedia

Convex or conical above, red or yellow surrounded by a pale ring, and diameter 3.0–6.0 mm; before maturity there is no apparent opening and at maturity bracts split at apex into a jagged opening; below slightly convex and paler than leaf with a small closed nipple which may or may not open.